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Several kinds of insects
and mites cause curious swellings on plants called galls. Some of
these growths become as large as a baseball and are smooth in
appearance. Others may be smaller than the head of a pin. Some have
very strange shapes with spines and knobs on them. They are common
on oak, willow, poplar, rose and many other plants grown in
California. The native oaks support more than 90 types of galls. No
part of an oak is free from infestation . . . galls may be found on
leaves, flowers, buds, twigs, branches, roots and even acorns.
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The most common gall inducers are tiny, dark wasps called cynipid or gall wasps. Certain moths, beetles, flies and a few other insects and mites also form galls. Galls consist only of plant tissue. In most cases, normal plant cells have been stimulated to multiply at an unusually high rate by the activity of a gall inducer. To successfully form a gall (and thereby successfully reproduce) the insect or mite must deposit its eggs at a very precise moment in the plant’s growth cycle.
A gall wasp initiates the process by
piercing a selected plant part with its egg-laying device and
depositing an egg or eggs inside the plant tissue. Fluids deposited with the
egg or produced by the larva after hatching cause the plant cell
multiplication process to begin. The larva produces additional substances that
maintain and control cell division that determines the size and
shape of the developing gall. The larva develops within a
cavity inside the gall, feeding on material produced on the cavity
lining. At maturity, it transforms into a pupa, and later it becomes
an adult that chews its way out of the gall. By causing the plant to
form a gall, the gall inducer has made the plant provided food and shelter for its
offspring.
Each gall inducer forms a gall of a particular size, shape and color; no other species forms one quite like it. (Click here for more gall images.) Gall inducers are specific about the types of plants and the plant parts they attack. Some galls contain more than a single larva of the gall inducer, but usually each lives within its own cavity.
Most insect or mite caused galls in California are not harmful to the plant. Several cause a scorching or spotting of leaves and a few result in death of twigs they infest. In nearly all cases prevention of gall formation is exceedingly difficult and is not considered practical. For many insect and mite species that cause galls, a means of prevention or control is unknown.